寄生虫标准株的筛选、鉴定、保藏与应用专家共识

doi:10.12140/j.issn.1000-7423.2026.03.001

中国寄生虫学与寄生虫病杂志 ›› 2026, Vol. 44 ›› Issue (3): 309-319.doi: 10.12140/j.issn.1000-7423.2026.03.001

寄生虫标准株的筛选、鉴定、保藏与应用专家共识

寄生虫标准株的筛选、鉴定、保藏与应用专家共识撰写组

收稿日期:2026-03-02 修回日期:2026-05-25 出版日期:2026-06-30 发布日期:2026-06-12
通讯作者: *周晓农（ORCID：0000000314178427），男，博士，研究员，从事寄生虫病空间流行病学、血吸虫病流行病学、钉螺生物学、寄生虫与宿主间相互关系等研究。Email：zhouxn1@chinacdc.cn
作者简介:李石柱（ORCID：0000-0003-3172-5218），男，博士，研究员，从事寄生虫病病原生物学和流行病学研究。E-mail：lisz@chinacdc.cn
基金资助:
国家科技基础条件平台国家寄生虫资源库项目(NPRC-2019-194-30)

Expert consensus on screening, identification, preservation and application of standard parasite strains

Expert Consensus Writing Group on the Screening, Identification, Preservation and Application of Parasite Standard Strains, National Parasitic Resources Center

Received:2026-03-02 Revised:2026-05-25 Online:2026-06-30 Published:2026-06-12
Contact: *Email: zhouxn1@chinacdc.cn; lisz@chinacdc.cn
Supported by:
National Parasitic Resources Center and the Ministry of Science and Technology Fund(NPRC-2019-194-30)

摘要/Abstract

摘要：

寄生虫标准株是寄生虫学研究、寄生虫病防控与生物医药创新的基础资源。基于我国寄生虫标准株体系建立不完善、规范化管理不足等问题，国家寄生虫资源库秘书处组织全国54位专家开展系统性研讨后达成重要共识。共识系统阐述了寄生虫标准株筛选、鉴定、保藏与应用的规范及细则，明确了标准株应具备代表性、遗传稳定性、纯度单一性，可通过形态学、分子生物学等多维技术进行鉴定与质量控制；同时，提出了分级保藏策略、标准化分发流程，以及细化在不同科研场景下的应用规范，并强调伦理审查、生物安全与资源惠益共享原则的实施要求。共识还提出了加强顶层设计、构建全链条保藏体系、推动人工智能技术应用、建立多学科协作平台的建议，以加速建成与国际接轨的中国寄生虫标准株体系，提升国家生物资源战略储备能力与科技创新竞争力。

关键词: 寄生虫, 标准株, 鉴定, 应用, 共识

Abstract:

Parasite standard strains are fundamental resources for parasitological research, disease control, and biopharmaceutical innovation. To address the incomplete system and lagging standardization of parasite strains in China, the National Parasite Resource Center organized 54 experts from all over the country to reach an important consensus after discussion, and the consensus system expounded the standardized rules for the screening, identification, preservation and application of parasite standard strains. The consensus clarified that parasite standard strains should have representativeness, genetic stability, and purity unity, which can be identified and quality controlled through multi-dimensional technologies such as morphology and molecular biology. Standardize distribution processes and application specifications in different scientific research scenarios, emphasizing the importance of ethical review, biosecurity, and resource benefit sharing. Suggestions are put forward to strengthen top-level design, improve the preservation system, promote the application of artificial intelligence (AI) technology, and establish a multidisciplinary collaboration platform to accelerate the construction of China’s parasite reference strain system in line with international standards, and enhance the national biological resource reserve capacity and scientific research competitiveness.

Key words: Parasite, Standard strain, Identification, Application, Consensus

中图分类号:

寄生虫标准株的筛选、鉴定、保藏与应用专家共识撰写组. 寄生虫标准株的筛选、鉴定、保藏与应用专家共识[J]. 中国寄生虫学与寄生虫病杂志, 2026, 44(3): 309-319.

Expert Consensus Writing Group on the Screening, Identification, Preservation and Application of Parasite Standard Strains, National Parasitic Resources Center. Expert consensus on screening, identification, preservation and application of standard parasite strains[J]. Chinese Journal of Parasitology and Parasitic Diseases, 2026, 44(3): 309-319.

参考文献 74

[1]	Campbell TA, VerCauteren KC. Diseases and parasites[M]//Hewitt DG. Biology and management of white-tailed deer. Boca Raton: CRC Press, 2011: 232-263.
[2]	Dobson A, Kutz S, Pascual M, et al. Pathogens and parasites in a changing climate[M]//Hannah L, Lovejoy TE. Climate change and biodiversity:synergistic impacts. New Haven: Yale University Press, 2003: 33-38.
[3]	Tompkins DM, Poulin R. Parasites and biological invasions[M]// AllenRB, LeeWG. Biologicalinvasions in New Zealand. Berlin/Heidelberg: Springer, 2006: 67-84.
[4]	Schmid-Hempel P. Sociality and parasite transmission[J]. Behav Ecol Sociobiol, 2021, 75(11): 156.
[5]	Karvonen A, Jokela J, Laine AL. Importance of sequence and timing in parasite coinfections[J]. Trends Parasitol, 2019, 35(2): 109-118.
[6]	de Couvreur LA, Cobo MJ, Kennedy PJ, et al. Bibliometric analysis of parasite vaccine research from 1990 to 2019[J]. Vaccine, 2023, 41(44): 6468-6477.
[7]	Elsheikha HM, Marra CM, Zhu XQ. Epidemiology, pathophysiology, diagnosis, and management of cerebral toxoplasmosis[J]. Clin Microbiol Rev, 2020, 34: e00115-e00119.
[8]	Kyrpides NC, Woyke T, Eisen JA, et al. Genomic encyclopedia of type strains, phase I: the one thousand microbial genomes (KMG-I) project[J]. Stand Genomic Sci, 2014, 9(3): 1278-1284.
[9]	H?rdt A, López MG, Meier-Kolthoff JP, et al. Analysis of 1,000+ type-strain genomes substantially improves taxonomic classification of Alphaproteobacteria[J]. Front Microbiol, 2020, 11: 468.
[10]	Tambo E, Khayeka-Wandabwa C, Muchiri GW, et al. China’s Belt and Road Initiative: incorporating public health measures toward global economic growth and shared prosperity[J]. Glob Health J, 2019, 3(2): 46-49.
[11]	Qian MB, Abela-Ridder B, Wu WP, et al. Combating echinococcosis in China: strengthening the research and development[J]. Infect Dis Poverty, 2017, 6: 161.
[12]	Boundy-Mills K, Hess M, Bennett AR, et al. The United States culture collection network (USCCN): enhancing microbial genomics research through living microbe culture collections[J]. Appl Environ Microbiol, 2015, 81(17): 5671-5674.
[13]	Clark WA, Geary DH. The story of the American Type Culture Collection: its history and development (1899-1973)[J]. Adv Appl Microbiol, 1974, 17: 295-309.
[14]	Molestina RE, Stedman TT. Update on BEI resources for parasitology and arthropod vector research[J]. Trends Parasitol, 2020, 36(4): 321-324.
[15]	Molestina RE. BEI Resources: a biological resource center for parasitologists[J]. Trends Parasitol, 2010, 26(12): 559-560.
[16]	Adams JH, Wu Y, Fairfield A. Malaria research and reference reagent resource center[J]. Parasitol Today, 2000, 16(3): 89.
[17]	Smith D. Culture collections over the world[J]. Int Microbiol, 2003, 6(2): 95-100.
[18]	Fesser A, Beilstein S, Kaiser M, et al. Trypanosoma cruzi STIB980: a TcI strain for drug discovery and reverse genetics[J]. Pathogens, 2023, 12(10): 1217.
[19]	Thiam R, Ceballos MS, Beneke T, et al. A novel Leishmania infantum reference strain for gene editing and the study of visceral leishmaniasis[J]. PLoS One, 2025, 20(9): e0327390.
[20]	Dijkshoorn L, Ursing BM, Ursing JB. Strain, clone and species: comments on three basic concepts of bacteriology[J]. J Med Microbiol, 2000, 49(5): 397-401.
[21]	Smith D. Culture collections[M]//Advances in applied microbiology Volume 79. Amsterdam: Elsevier, 2012: 73-118.
[22]	Manning FJ, Bond EC, Berns KI. Resource Sharing in Biomedical Research[M]. Washington, D.C.: National Academies Press, 1996.
[23]	Fux CA, Shirtliff M, Stoodley P, et al. Can laboratory reference strains mirror ‘real-world’ pathogenesis?[J]. Trends Microbiol, 2005, 13(2): 58-63.
[24]	Schuster FL. Cultivation of Plasmodium spp[J]. Clin Microbiol Rev, 2002, 15(3): 355-364.
[25]	Dorman MJ, Thomson NR. ‘Community evolution’ - laboratory strains and pedigrees in the age of genomics[J]. Microbiology (Reading), 2020, 166(3): 233-238.
[26]	Adika E, Saiid S, Amoako EK, et al. Plasmodium falciparum isolates: ex vivo drug response[J]. J Antimicrob Chemother, 2025, 80(7): 1813-1822.
[27]	Habtewold T, Lwetoijera DW, Hoermann A, et al. Gene-drive-capable mosquitoes suppress patient-derived malaria in Tanzania[J]. Nature, 2026, 649(8096): 442-448.
[28]	Estelle DL, Jacques GE, Issiaka S, et al. Unstable laboratory Wolbachia strain w-Anga is negatively correlated with Plasmodium falciparum in wild malaria vectors[J]. Sci Rep, 2025, 15(1): 17732.
[29]	Gould SJ, Johnston RF. Geographic variation[J]. Annu Rev Ecol Syst, 1972, 3 (1972): 457-498.
[30]	Rocaboy C, Dardé ML, Aubert D, et al. Biological resource center for Toxoplasma[J]. Open J Bioresour, 2020, 7: 7.
[31]	Van Rossum T, Ferretti P, Maistrenko OM, et al. Diversity within species: interpreting strains in microbiomes[J]. Nat Rev Microbiol, 2020, 18(9): 491-506.
[32]	Miyake Y, Karanis P, Uga S. Cryopreservation of protozoan parasites[J]. Cryobiology, 2004, 48(1): 1-7.
[33]	Miyata A. Cryo-preservation of the parasitic protozoa[J]. Jap J Trop Med Hyg, 1975, 3(2): 161-200.
[34]	Frank SA. Models of parasite virulence[J]. Q Rev Biol, 1996, 71(1): 37-78.
[35]	Dougherty ER, Carlson CJ, Bueno VM, et al. Paradigms for parasite conservation[J]. Conserv Biol, 2016, 30(4): 724-733.
[36]	Forneris NS, Legarra A, Vitezica ZG, et al. Quality control of genotypes using heritability estimates of gene content at the marker[J]. Genetics, 2015, 199(3): 675-681.
[37]	Benavides F, Rülicke T, Prins JB, et al. Genetic quality assurance and genetic monitoring of laboratory mice and rats: FELASA Working Group Report[J]. Lab Anim, 2020, 54(2): 135-148.
[38]	Unemo M, Golparian D, Sánchez-Busó L, et al. The novel 2016 WHO Neisseria gonorrhoeae reference strains for global quality assurance of laboratory investigations: phenotypic, genetic and reference genome characterization[J]. J Antimicrob Chemother, 2016, 71(11): 3096-3108.
[39]	Sousa S, Fernandes M, Correia da Costa JM. Serotyping, a challenging approach for Toxoplasma gondii typing[J]. Front Med, 2023, 10: 1111509.
[40]	Pal C. Redox modulating small molecules having antimalarial efficacy[J]. Biochem Pharmacol, 2023, 218: 115927.
[41]	Batista FA, Gyau B, Vilacha JF, et al. New directions in antimalarial target validation[J]. Expert Opin Drug Discov, 2020, 15(2): 189-202.
[42]	Mostafa El-Sayed N, Gamal Masoud N. Medicinal plants as natural anti-parasitic agents against Blastocystis species[J]. Recent Adv Antiinfect Drug Discov, 2023, 18(1): 2-15.
[43]	Stanisic DI, McCarthy JS, Good MF. Controlled human malaria infection: applications, advances, and challenges[J]. Infect Immun, 2018, 86(1): e00479-e00417.
[44]	Gallichotte EN, Dobos KM, Ebel GD, et al. Towards a method for cryopreservation of mosquito vectors of human pathogens[J]. Cryobiology, 2021, 99: 1-10.
[45]	Unemo M, Fasth O, Fredlund H, et al. Phenotypic and genetic characterization of the 2008 WHO Neisseria gonorrhoeae reference strain panel intended for global quality assurance and quality control of gonococcal antimicrobial resistance surveillance for public health purposes[J]. J Antimicrob Chemother, 2009, 63(6): 1142-1151.
[46]	Gugosyan YA, Boyko OO, Brygadyrenko VV. Morphological variation of four species of Strongyloides (Nematoda, Rhabditida) parasitising various mammal species[J]. Biosys Divers, 2019, 27(1): 85-98.
[47]	Brazenor AK, Saunders RJ, Miller TL, et al. Morphological variation in the cosmopolitan fish parasite Neobenedenia girellae (Capsalidae∶Monogenea)[J]. Int J Parasitol, 2018, 48(2): 125-134.
[48]	Veiga J, De O?a P, Salazar B, et al. Defining host range: host-parasite compatibility during the non-infective phase of the parasite also matters[J]. Parasitology, 2019, 146(2): 234-240.
[49]	Heath MC. A generalized concept of host-parasite specificity[J]. Phytopathology, 1981, 71(11): 1121.
[50]	Marcogliese DJ. Parasites of the superorganism: are they indicators of ecosystem health?[J]. Int J Parasitol, 2005, 35(7): 705-716.
[51]	Biswal D. Fish parasites as biological indicators: a systematic review[J]. Biosci Biotech Res Comm, 2020, 13(4): 1743-1755.
[52]	Combes C. Ethological aspects of parasite transmission[J]. Am Nat, 1991, 138(4): 866-880.
[53]	Fenton A, Fairbairn JP, Norman R, et al. Parasite transmission: reconciling theory and reality[J]. J Anim Ecol, 2002, 71(5): 893-905.
[54]	Alvarez JA, Rojas C, Figueroa JV. An overview of current knowledge on in vitro Babesia cultivation for production of live attenuated vaccines for bovine babesiosis in Mexico[J]. Front Vet Sci, 2020, 7: 364.
[55]	Stanisic DI, Good MF. Malaria vaccines: progress to date[J]. BioDrugs, 2023, 37(6): 737-756.
[56]	Sonstegard TS, Gasbarre LC. Genomic tools to improve parasite resistance[J]. Vet Parasitol, 2001, 101(3/4): 387-403.
[57]	Duflot M, Midelet G, Bourgau O, et al. Optimization of tools for the detection and identification of Cryptocotyle metacercariae in fish: digestion method and viability studies[J]. J Fish Dis, 2021, 44(11): 1777-1784.
[58]	Van Wyk JA, Mayhew E. Morphological identification of parasitic nematode infective larvae of small ruminants and cattle: a practical lab guide[J]. Onderstepoort J Vet Res, 2013, 80(1): 1-14.
[59]	Nugroho HA, Ali Akbar S, Murhandarwati EEH. Feature extraction and classification for detection malaria parasites in thin blood smear[C]// 2015 2nd International Conference on Information Technology, Computer, and Electrical Engineering (ICITACEE). Semarang, Indonesia. IEEE, 2015: 197-201.
[60]	潘严, 姜子睿, 周超, 等. 肠道寄生虫粪便的病原学检查方法[J]. 中国医药科学, 2016, 6(20): 213-217.
	Pan Y, Jiang ZR, Zhou C, et al. Pathogenic examination methods of intestinal parasite stool[J]. China Med Pharm, 2016, 6(20): 213-217. (in Chinese)
[61]	何婷. 分泌物涂片检查霉菌及滴虫性阴道炎的结果研究[J]. 系统医学, 2022, 7(19): 91-94.
	He T. Results of secretion smear examination of colpomycosis and trichomonal vaginitis[J]. Syst Med, 2022, 7(19): 91-94. (in Chinese)
[62]	张时民, 张峰. 痰液中检出肺吸虫卵致肺吸虫感染一例[J]. 中华检验医学杂志, 2011, 34(8): 755-756.
	Zhang SM, Zhang F. A case of Paragonimus infection caused by detection of Paragonimus eggs in sputum[J]. Chin J Lab Med, 2011, 34(8): 755-756. (in Chinese)
[63]	杨绮红, 舒建昌, 黎铭恩, 等. 十二指肠胆汁引流术诊断隐匿型华支睾吸虫感染[J]. 中国热带医学, 2009, 9(12): 2244, 2271.
	Yang QH, Shu JC, Li ME, et al. Diagnosis of Clonorchis sinensis infection by duodenum biliary drainage aided with gastroscope[J]. China Trop Med, 2009, 9(12): 2244, 2271. (in Chinese)
[64]	林爱芬. 对经肋间、腹部的脾穿刺术及骨髓穿刺术用于诊断内脏利什曼病的比较研究[J]. 国外医学(寄生虫病分册), 1999, 26(2): 93.
	Lin AF. A comparative study of intercostal, abdominal splenic puncture and bone marrow puncture for the diagnosis of visceral leishmaniasis[J]. Int J Med Parasit Dis, 1999, 26(2): 93. (in Chinese)
[65]	Palanisamy A, Gochhait D, Sekar MD, et al. A case of cutaneous leishmaniasis: an emphasis on scrape cytology as a diagnostic modality[J]. J Cytol, 2025, 42(2): 101-103.
[66]	张秉西, 高晓娜. 压片法显微镜诊断肠内寄生虫的临床意义[J]. 现代医药卫生, 2004, 20(21): 2295.
	Zhang BX, Gao XN. Clinical significance of microscopic diagnosis of intestinal parasites by compression method[J]. J Mod Med Heal, 2004, 20(21): 2295. (in Chinese)
[67]	柳志成, 车金, 柴宜均, 等. 牛血液寄生虫耐药性研究进展[J]. 中国草食动物科学, 2025, 45(3): 87-91.
	Liu ZC, Che J, Chai YJ, et al. Advances in drug resistance of bovine blood parasites[J]. China Herbiv Sci, 2025, 45(3): 87-91. (in Chinese)
[68]	吕照勇, 王锦明, 王晓星, 等. 羊巴贝斯虫SBP3基因序列分析和诊断标识潜力评价[J]. 中国兽医科学, 2021, 51(7): 840-847.
	Lü ZY, Wang JM, Wang XX, et al. Sequence analysis of ovine Babesia SBP3 gene and evaluation of potential as a diagnostic marker[J]. Chin Vet Sci, 2021, 51(7): 840-847. (in Chinese)
[69]	孙艾明, 董莹, 陈梦妮, 等. 云南省恶性疟原虫青蒿素耐药性相关基因K13 kelch结构域序列多态性的分析[J]. 中国寄生虫学与寄生虫病杂志, 2016, 34(4): 339-345.
	Sun AM, Dong Y, Chen MN, et al. Polymorphism analysis of Plasmodium falciparum K13 gene kelch domain associated with resistance to artemisinin in Yunnan Province[J]. Chin J Parasitol Parasit Dis, 2016, 34(4): 339-345. (in Chinese)
[70]	Lu Y, Zhang D, Han DS, et al. Case report: diagnosis of hemolytic anemia from Babesia and secondary multi-pathogen pneumonia using a metagenomic next-generation sequencing approach[J]. Infect Drug Resist, 2024, 17: 3785-3791.
[71]	Singh P, Lonardi S, Liang QH, et al. Babesia duncani multi-omics identifies virulence factors and drug targets[J]. Nat Microbiol, 2023, 8(5): 845-859.
[72]	段磊, 屈磊, 郭云海, 等. 我国藁杆双脐螺分子溯源研究[J]. 中国血吸虫病防治杂志, 2024, 36(3): 272-278, 285.
	Duan L, Qu L, Guo YH, et al. Molecular tracing of Biomphalaria straminea in China[J]. Chin J Schistosomiasis Control, 2024, 36(3): 272-278, 285. (in Chinese)
[73]	Alaithan H, Kumar N, Islam MZ, et al. Novel therapeutics for malaria[J]. Pharmaceutics, 2023, 15(7): 1800.
[74]	国家卫生健康委员会. 病原微生物菌(毒)种国家标准株评价技术标准: WS/T 812—2022[S]. 北京: 中国标准出版社, 2022.
	National Health Commission of the People’s Republic of China. Technical requirements for evaluation of national standard strains of pathogenic microorganisms: WS/T 812—2022[S]. Beijing: Standards Press of China, 2022. (in Chinese)

寄生虫标准株的筛选、鉴定、保藏与应用专家共识

Expert consensus on screening, identification, preservation and application of standard parasite strains

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[12]	秦若芳, 许海萍, 胥奕炯, 戴思敏, 吕山, 崔燕. 2017—2024年上海市普陀区流通食品食源性寄生虫监测分析[J]. 中国寄生虫学与寄生虫病杂志, 2025, 43(5): 690-696.
[13]	许晓娟, 陈雪峰, 吴凡, 吴晨阳, 刘婷, 代波, 张世清. 2022年安徽省沿江地区野鼠肠道寄生虫感染情况[J]. 中国寄生虫学与寄生虫病杂志, 2025, 43(5): 727-730.
[14]	朱民, 朱国鼎, 王笑笑, 许娴, 王真瑜, 曹园园, 丰燕, 张滔, 张宸罡, 张轩, 余晴. 长江三角洲输入性疟疾联合防控技术措施——沪苏浙皖专家共识[J]. 中国寄生虫学与寄生虫病杂志, 2025, 43(4): 458-462.
[15]	王旭, 尹建海, 韩帅, 杨扬, 李梦晴, 沈玉娟, 曹建平. 寄生虫的健康威胁与生态功能[J]. 中国寄生虫学与寄生虫病杂志, 2025, 43(4): 463-468.