中国寄生虫学与寄生虫病杂志 ›› 2004, Vol. 22 ›› Issue (2): 14-118.

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无前螨科沙螨 (恙螨 )的分类(英文)

温廷桓
  

  1. 复旦大学上海医学院医学螨类研究室,上海 200032
  • 收稿日期:1900-01-01 修回日期:1900-01-01 出版日期:2004-04-30 发布日期:2004-04-30

Classification of the Sand-mite Family Walchiidae (Acariformes: Trombiculoidea)

WEN Ting-huan
  

  1. Medical Acarology Laboratory,shanghai Medical College,Fudan University,Shanghai 200032,China
  • Received:1900-01-01 Revised:1900-01-01 Online:2004-04-30 Published:2004-04-30

摘要:   目的 无前螨亚科WalchiinaeEwing,1946与甲逦螨亚科GahrliepiinaeWomersley,1952两者孰为符合动物命名法规的优先名,争论自1952年起经有半个世纪。无前螨亚科的建立是以无前螨属WalchiaEwing ,1931为模式属在先 ,以后甲逦螨亚科的建立是以甲逦螨属GahrliepiaOudemans,1912为模式属 ,将无前螨属置于其亚属。现两属均为有效属。作者(1999)对于该两亚科赋予各自新的特征 ,并提升无前螨亚科为科级阶元 ,无前螨科Walchiidae (Ewing,1946)Wen,1999。无前螨科的特征为SIF=4B/4Bs/5B/6BN/B3/22(1)1(0)1(0)0.0000,螨体小至大型,IP=320~1220。盾板小至大型 ,在背面向后伸展 ,五角形后端尖锐或呈舌形。不具前中毛 (AM或vi)和前中突(A或N=0),盾板基本毛式为AL和PL各一对,常有附加毛PPLs2~40支不等,少数还有间毛(IM)1~2对。感器(Sn或sci)全为短棒杆形。足节式fSP=7.6.6恒定。盾板AM的退化、足节的增多、体触觉毛的减少是沙螨(恙螨)的衍征,意味无前螨科比沙螨科(恙螨科)Trombiculidae(Ewing,1929)和列螨科Leeuwenhoekiidae (Womersley,1944)进化,三科综合为沙螨总科Trombiculoidea(Ewing,1929)necWelbourn,1991sensuWen,1999,可使脊椎动物

关键词: 分类法, 无前螨科, 沙螨(恙螨)总科

Abstract:  The priority of Walchiinae Ewing, 1946 based on Walchia Ewing, 1931 or Gahrliepiinae Womersley, 1952 based on Gahrliepia Oudemans, 1912 as Walchia regarded as subgeneric taxon had been a controversy for almost half a century since 1952. Both genera are valid now. Wen (1999) redefined both subfamilial characters and in turn promoted Walchiinae to a full (fami)lial status. Walchiidae (Ewing,1946) Wen, 1999 is characterized by SIF=4B/4Bs/5B/6B-N/B-3/2-2(1)1(0)1(0)0.0000, small to large sized sand-mites, IP=320-1220. Scutum is small to large size, extending backward over part of dorsum, and pentagonal with acuminate posterior angle or tongue-shaped. The scutum is never provided with anteromedian setae (AM or vi) and anteromedian projection(A or N=0). Scutal setae have AL and PL pairs basically, frequently in addition with 2-40 accessories (PPLs), and rarely 1-2 pairs of intermedial setae (IM). Sensillae (Sn or sci) are short and expanded. Leg segments are 7.6.6 always without variations.Casting off the anteromedian setae on scutum, increasing the leg segments and reducing the tactile body setae are the plesiomorphic characters of sand-mites, that means Walchiidae in higher advance of evolution than both Trombiculidae (Ewing, 1929) and Leeuwenhoekiidae (Womersley,1944). It is rationally to unify three families of vertebrate parasitic larvae into a single superfamily, Trombiculoidea nec Welbourn (1991), that separable from superfamily Trombidioidea of arthropod parasitic larvae and standing at most advanced evolution of Parasitengona.Family Walchiidae has 2 subfamilies, Walchiinae Ewing, 1946 sensu Wen 1999 and Gahrliepiinae Womersley, 1952, sensu Wen 1999. Each subfamily contains two tribes, Walchiini (Ewing, 1946) Wen 1984 and Schoengastiellini Wen 1984 for Walchiinae, and Gahrliepiini sensu Wen 1984 and Intermedialiini Wen, 1984 for Gahrliepiinae. Currently this family has 18 genera and 28 subgenera, 248 nominated species and subspecies. Walchiid sand-mites are essentially an Old World family and best developed in the Oriental Region with the center of development in Southeast Asia.

Key words: classification, Walchiidae, Trombiculoidea